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Professor Tim Coulson

Publications


Journal Articles

  • Wallace KM, Leslie AJ, Coulson T, Living with predators: a focus on the issues of human-crocodile conflict within the lower Zambezi valley, WILDLIFE RES, 2011, Vol:38, Pages:747-755, ISSN:1035-3712(doi)
  • Coulson T, MacNulty DR, Stahler DR, et al, Modeling effects of environmental change on wolf population dynamics, trait evolution, and life history., Science, 2011, Vol:334, Pages:1275-1278(doi)
  • Bateman AW, Ozgul A, Coulson T, et al, Density dependence in group dynamics of a highly social mongoose, Suricata suricatta., J Anim Ecol, 2011(doi)
  • Nussey DH, Coulson T, Delorme D, et al, Patterns of body mass senescence and selective disappearance differ among three species of free-living ungulates., Ecology, 2011, Vol:92, Pages:1936-1947, ISSN:0012-9658
  • Pettorelli N, Coulson T, Durant SM, et al, Predation, individual variability and vertebrate population dynamics., Oecologia, 2011, Vol:167, Pages:305-314(doi)
  • Childs DZ, Coulson TN, Pemberton JM, et al, Predicting trait values and measuring selection in complex life histories: reproductive allocation decisions in Soay sheep., Ecol Lett, 2011, Vol:14, Pages:985-992(doi)
  • Pelletier F, Moyes K, Clutton-Brock TH, et al, Decomposing variation in population growth into contributions from environment and phenotypes in an age-structured population., Proc Biol Sci, 2012, Vol:279, Pages:394-401(doi)
  • Smallegange IM, Coulson T, The stochastic demography of two coexisting male morphs., Ecology, 2011, Vol:92, Pages:755-764, ISSN:0012-9658
  • Roy BA, Coulson T, Blaser W, et al, Population regulation by enemies of the grass Brachypodium sylvaticum: demography in native and invaded ranges., Ecology, 2011, Vol:92, Pages:665-675, ISSN:0012-9658
  • Di Fonzo MM, Pelletier F, Clutton-Brock TH, et al, The population growth consequences of variation in individual heterozygosity., PLoS One, 2011, Vol:6(doi)
  • Bateman AW, Coulson T, Clutton-Brock TH, What do simple models reveal about the population dynamics of a cooperatively breeding species?, OIKOS, 2011, Vol:120, Pages:787-794, ISSN:0030-1299(doi)
  • Moyes K, Morgan B, Morris A, et al, Individual differences in reproductive costs examined using multi-state methods., J Anim Ecol, 2011, Vol:80, Pages:456-465(doi)
  • Horvitz CC, Coulson T, Tuljapurkar S, et al, A NEW WAY TO INTEGRATE SELECTION WHEN BOTH DEMOGRAPHY AND SELECTION GRADIENTS VARY OVER TIME, INT J PLANT SCI, 2010, Vol:171, Pages:945-959, ISSN:1058-5893(doi)
  • Simard MA, Coulson T, Gingras A, et al, Influence of Density and Climate on Population Dynamics of a Large Herbivore Under Harsh Environmental Conditions, J WILDLIFE MANAGE, 2010, Vol:74, Pages:1671-1685, ISSN:0022-541X(doi)
  • Coulson T, Tuljapurkar S, Childs DZ, Using evolutionary demography to link life history theory, quantitative genetics and population ecology., J Anim Ecol, 2010, Vol:79, Pages:1226-1240(doi)
  • Ozgul A, Childs DZ, Oli MK, et al, Coupled dynamics of body mass and population growth in response to environmental change., Nature, 2010, Vol:466, Pages:482-485(doi)
  • Hart T, Mann R, Coulson T, et al, Behavioural switching in a central place forager: patterns of diving behaviour in the macaroni penguin (Eudyptes chrysolophus), MAR BIOL, 2010, Vol:157, Pages:1543-1553, ISSN:0025-3162(doi)
  • Smallegange IM, Coulson T, Unifying ecological and evolutionary dynamics through experimental stochastic demography., Israel Journal of Ecology & Evolution, 2009, Vol:55, Pages:199-205
  • Rebke M, Coulson T, Becker PH, et al, Reproductive improvement and senescence in a long-lived bird., Proc Natl Acad Sci U S A, 2010, Vol:107, Pages:7841-7846(doi)
  • Hart T, Coulson T, Trathan PN, Time series analysis of biologging data: autocorrelation reveals periodicity of diving behaviour in macaroni penguins, ANIM BEHAV, 2010, Vol:79, Pages:845-855, ISSN:0003-3472(doi)
  • Ezard THG, Coulson T, How sensitive are elasticities of long-run stochastic growth to how environmental variability is modelled?, ECOL MODEL, 2010, Vol:221, Pages:191-200, ISSN:0304-3800(doi)
  • Aust P, Boyle B, Fergusson R, et al, The impact of Nile crocodiles on rural livelihoods in northeastern Namibia, S AFR J WILDL RES, 2009, Vol:39, Pages:57-69, ISSN:0379-4369
  • Hart T, Fitzcharles E, Trathan PN, et al, Testing and Improving the Accuracy of Discriminant Function Tests: A Comparison between Morphometric and Molecular Sexing in Macaroni Penguins, WATERBIRDS, 2009, Vol:32, Pages:437-443, ISSN:1524-4695
  • Bonenfant C, Gaillard JM, Coulson T, et al, Empirical Evidence of Density-Dependence in Populations of Large Herbivores, ADV ECOL RES, 2009, Vol:41, Pages:313-357, ISSN:0065-2504(doi)
  • Ozgul A, Tuljapurkar S, Benton TG, et al, The Dynamics of Phenotypic Change and the Shrinking Sheep of St. Kilda, SCIENCE, 2009, Vol:325, Pages:464-467, ISSN:0036-8075(doi)
  • Haridas CV, Tuljapurkar S, Coulson T, Estimating stochastic elasticities directly from longitudinal data, ECOL LETT, 2009, Vol:12, Pages:806-812, ISSN:1461-023X(doi)
  • Martinez-Jauregui M, San Miguel-Ayanz A, Mysterud A, et al, Are local weather, NDVI and NAO consistent determinants of red deer weight across three contrasting European countries?, GLOBAL CHANGE BIOL, 2009, Vol:15, Pages:1727-1738, ISSN:1354-1013(doi)
  • Malo AF, Coulson T, Heterozygosity-fitness correlations and associative overdominance: new detection method and proof of principle in the Iberian wild boar., Mol Ecol, 2009, Vol:18, Pages:2741-2742(doi)
  • Tavecchia G, Besbeas P, Coulson T, et al, Estimating Population Size and Hidden Demographic Parameters with State-Space Modeling, AM NAT, 2009, Vol:173, Pages:722-733, ISSN:0003-0147(doi)
  • Tuljapurkar S, Gaillard JM, Coulson T, From stochastic environments to life histories and back, PHILOS T R SOC B, 2009, Vol:364, Pages:1499-1509, ISSN:0962-8436(doi)
  • Coulson T, 'A review of extinction in experimental populations' by Blaine Griffen and John Drake., J Anim Ecol, 2008, Vol:77(doi)
  • Da Silva A, Gaillard JM, Yoccoz NG, et al, Heterozygosity-fitness correlations revealed by neutral and candidate gene markers in roe deer from a long-term study., Evolution, 2009, Vol:63, Pages:403-417(doi)
  • McLoughlin PD, Coulson T, Clutton-Brock T, Cross-generational effects of habitat and density on life history in red deer., Ecology, 2008, Vol:89, Pages:3317-3326, ISSN:0012-9658
  • McLoughlin PD, Coulson T, Clutton-Brock T, CROSS-GENERATIONAL EFFECTS OF HABITAT AND DENSITY ON LIFE HISTORY IN RED DEER, ECOLOGY, 2008, Vol:89, Pages:3317-3326, ISSN:0012-9658
  • King R, Brooks SP, Coulson T, Analyzing Complex Capture-Recapture Data in the Presence of Individual and Temporal Covariates and Model Uncertainty, BIOMETRICS, 2008, Vol:64, Pages:1187-1195, ISSN:0006-341X(doi)
  • Moyes K, Morgan BJ, Morris A, et al, Exploring individual quality in a wild population of red deer., J Anim Ecol, 2009, Vol:78, Pages:406-413(doi)
  • Coulson T, Tuljapurkar S, The Dynamics of a Quantitative Trait in an Age-Structured Population Living in a Variable Environment, AM NAT, 2008, Vol:172, Pages:599-612, ISSN:0003-0147(doi)
  • Coulson T, Malo A, Population biology: Case of the absent lemmings, NATURE, 2008, Vol:456, Pages:43-44, ISSN:0028-0836(doi)
  • Proaktor G, Coulson T, Milner-Gulland EJ, The demographic consequences of the cost of reproduction in ungulates, ECOLOGY, 2008, Vol:89, Pages:2604-2611, ISSN:0012-9658
  • Kimber J, Hickman M, Degenhardt L, et al, Estimating the size and dynamics of an injecting drug user population and implications for health service coverage: comparison of indirect prevalence estimation methods, ADDICTION, 2008, Vol:103, Pages:1604-1613, ISSN:0965-2140(doi)
  • Stopher KV, Pemberton JM, Clutton-Brock TH, et al, Individual differences, density dependence and offspring birth traits in a population of red deer, P R SOC B, 2008, Vol:275, Pages:2137-2145, ISSN:0962-8452(doi)
  • Ezard TH, Gaillard JM, Crawley MJ, et al, Habitat dependence and correlations between elasticities of long-term growth rates., Am Nat, 2008, Vol:172, Pages:424-430(doi)
  • Coulson T, Ezard THG, Pelletier F, et al, Estimating the functional form for the density dependence from life history data, ECOLOGY, 2008, Vol:89, Pages:1661-1674, ISSN:0012-9658
  • Jones OR, Gaillard JM, Tuljapurkar S, et al, Senescence rates are determined by ranking on the fast-slow life-history continuum, ECOL LETT, 2008, Vol:11, Pages:664-673, ISSN:1461-023X(doi)
  • Nussey DH, Coulson T, Festa-Bianchet M, et al, Measuring senescence in wild animal populations: towards a longitudinal approach, FUNCT ECOL, 2008, Vol:22, Pages:393-406, ISSN:0269-8463(doi)
  • Jones OR, Clutton-Brock T, Coulson T, et al, A web resource for the UK's long-term individual-based time-series (LITS) data., J Anim Ecol, 2008, Vol:77, Pages:612-615(doi)
  • Sandvik H, Coulson T, Saether BE, A latitudinal gradient in climate effects on seabird demography: results from interspecific analyses, GLOBAL CHANGE BIOL, 2008, Vol:14, Pages:703-713, ISSN:1354-1013(doi)
  • Morris WF, Pfister CA, Tuljapurkar S, et al, Longevity can buffer plant and animal populations against changing climatic variability., Ecology, 2008, Vol:89, Pages:19-25, ISSN:0012-9658
  • Proaktor G, Milner-Gulland EJ, Coulson T, Age-related shapes of the cost of reproduction in vertebrates., Biol Lett, 2007, Vol:3, Pages:674-677, ISSN:1744-9561(doi)
  • Coulson T, Population ecology: group living and hungry lions., Nature, 2007, Vol:449, Pages:996-997(doi)
  • Ezard THG, Becker PH, Coulson T, Correlations between age, phenotype, and individual contribution to population growth in common terns, ECOLOGY, 2007, Vol:88, Pages:2496-2504, ISSN:0012-9658
  • Proaktor G, Coulson T, Milner-Gulland EJ, Evolutionary responses to harvesting in ungulates, J ANIM ECOL, 2007, Vol:76, Pages:669-678, ISSN:0021-8790(doi)
  • Foerster K, Coulson T, Sheldon BC, et al, Sexually antagonistic genetic variation for fitness in red deer, NATURE, 2007, Vol:447, Pages:1107-U9, ISSN:0028-0836(doi)
  • Nilsen EB, Milner-Gulland EJ, Schofield L, et al, Wolf reintroduction to Scotland: public attitudes and consequences for red deer management., Proc Biol Sci, 2007, Vol:274, Pages:995-1002, ISSN:0962-8452(doi)
  • Pelletier F, Clutton-Brock T, Pemberton J, et al, The evolutionary demography of ecological change: linking trait variation and population growth., Science, 2007, Vol:315, Pages:1571-1574(doi)
  • Moyes K, Coulson T, Morgan BJT, et al, Cumulative reproduction and survival costs in female red deer, OIKOS, 2006, Vol:115, Pages:241-252, ISSN:0030-1299
  • Borger L, Franconi N, De Michele G, et al, Effects of sampling regime on the mean and variance of home range size estimates, J ANIM ECOL, 2006, Vol:75, Pages:1393-1405, ISSN:0021-8790(doi)
  • Coulson T, Benton TG, Lundberg P, et al, Putting evolutionary biology back in the ecological theatre: a demographic framework mapping genes to communities, EVOL ECOL RES, 2006, Vol:8, Pages:1155-1171, ISSN:1522-0613
  • Ezard THG, Becker PH, Coulson T, The contributions of age and sex to variation in common tern population growth rate, J ANIM ECOL, 2006, Vol:75, Pages:1379-1386, ISSN:0021-8790(doi)
  • Börger L, Franconi N, Ferretti F, et al, An integrated approach to identify spatiotemporal and individual-level determinants of animal home range size., Am Nat, 2006, Vol:168, Pages:471-485(doi)
  • Lande R, Engen S, Saether BE, et al, Estimating density dependence from time series of population age structure, AM NAT, 2006, Vol:168, Pages:76-87, ISSN:0003-0147
  • McLoughlin PD, Boyce MS, Coulson T, et al, Lifetime reproductive success and density-dependent, multi-variable resource selection., Proc Biol Sci, 2006, Vol:273, Pages:1449-1454, ISSN:0962-8452(doi)
  • Festa-Bianchet M, Coulson T, Gaillard JM, et al, Stochastic predation events and population persistence in bighorn sheep., Proc Biol Sci, 2006, Vol:273, Pages:1537-1543, ISSN:0962-8452(doi)
  • Benton TG, Plaistow SJ, Coulson TN, Complex population dynamics and complex causation: devils, details and demography., Proc Biol Sci, 2006, Vol:273, Pages:1173-1181, ISSN:0962-8452(doi)
  • Coulson T, Benton TG, Lundberg P, et al, Estimating individual contributions to population growth: evolutionary fitness in ecological time, P ROY SOC B-BIOL SCI, 2006, Vol:273, Pages:547-555, ISSN:0962-8452(doi)
  • King R, Brooks SP, Morgan BJ, et al, Factors influencing soay sheep survival: a Bayesian analysis., Biometrics, 2006, Vol:62, Pages:211-220, ISSN:0006-341X(doi)
  • Jones JPG, Coulson T, Population regulation and demography in a harvested freshwater crayfish from Madagascar, OIKOS, 2006, Vol:112, Pages:602-611, ISSN:0030-1299
  • Holand O, Mysterud A, Roed KH, et al, Adaptive adjustment of offspring sex ratio and maternal reproductive effort in an iteroparous mammal, P R SOC B, 2006, Vol:273, Pages:293-299, ISSN:0962-8452(doi)
  • Nussey DH, Coltman DW, Coulson T, et al, Rapidly declining fine-scale spatial genetic structure in female red deer, MOL ECOL, 2005, Vol:14, Pages:3395-3405, ISSN:0962-1083
  • Martinez M, Rodriguez-Vigal C, Jones OR, et al, Different hunting strategies select for different weights in red deer, BIOL LETT-UK, 2005, Vol:1, Pages:353-356, ISSN:1744-9561(doi)
  • Coulson T, Gaillard JM, Festa-Bianchet M, Decomposing the variation in population growth into contributions from multiple demographic rates, J ANIM ECOL, 2005, Vol:74, Pages:789-801, ISSN:0021-8790(doi)
  • Ashenafi ZT, Coulson T, Sillero-Zubiri C, et al, Behaviour and ecology of the Ethiopian wolf (Canis simensis) in a human-dominated landscape outside protected areas, ANIM CONSERV, 2005, Vol:8, Pages:113-121, ISSN:1367-9430(doi)
  • Tavecchia G, Coulson T, Morgan BJT, et al, Predictors of reproductive cost in female Soay sheep, J ANIM ECOL, 2005, Vol:74, Pages:201-213, ISSN:0021-8790(doi)
  • Stenseth NC, Chan KS, Tavecchia G, et al, Modelling non-additive and nonlinear signals from climatic noise in ecological time series: Soay sheep as an example, P ROY SOC B-BIOL SCI, 2004, Vol:271, Pages:1985-1993, ISSN:0962-8452(doi)
  • Balloux F, Amos W, Coulson T, Does heterozygosity estimate inbreeding in real populations?, MOL ECOL, 2004, Vol:13, Pages:3021-3031, ISSN:0962-1083(doi)
  • Milner-Gulland EJ, Coulson T, Clutton-Brock TH, Sex differences and data quality as determinants of income from hunting red deer Cervus elaphus, WILDLIFE BIOL, 2004, Vol:10, Pages:187-201, ISSN:0909-6396
  • Bean K, Amos W, Pomeroy PP, et al, Patterns of parental relatedness and pup survival in the grey seal (Halichoerus grypus)., Mol Ecol, 2004, Vol:13, Pages:2365-2370, ISSN:0962-1083(doi)
  • Coulson T, Rohani P, Pascual M, Skeletons, noise and population growth: the end of an old debate?, TRENDS ECOL EVOL, 2004, Vol:19, Pages:359-364, ISSN:0169-5347(doi)
  • Hallett TB, Coulson T, Pilkington JG, et al, Why large-scale climate indices seem to predict ecological processes better than local weather, NATURE, 2004, Vol:430, Pages:71-75, ISSN:0028-0836(doi)
  • Clutton-Brock TH, Coulson T, Milner JM, Red deer stocks in the Highlands of Scotland, NATURE, 2004, Vol:429, Pages:261-262, ISSN:0028-0836(doi)
  • Catchpole EA, Fan Y, Morgan BJT, et al, Sexual dimorphism, survival and dispersal in red deer, J AGR BIOL ENVIR ST, 2004, Vol:9, Pages:1-26, ISSN:1085-7117(doi)
  • Coulson T, Guinness F, Pemberton J, et al, The demographic consequences of releasing a population of red deer from culling, ECOLOGY, 2004, Vol:85, Pages:411-422, ISSN:0012-9658
  • Catchpole EA, Morgan BJT, Coulson T, Conditional methodology for individual case history data, J ROY STAT SOC C-APP, 2004, Vol:53, Pages:123-131, ISSN:0035-9254
  • Coulson T, Kruuk LEB, Tavecchia G, et al, Estimating selection on neonatal traits in red deer using elasticity path analysis, EVOLUTION, 2003, Vol:57, Pages:2879-2892, ISSN:0014-3820
  • O'Brien S, Emahalala ER, Beard V, et al, Decline of the Madagascar radiated tortoise Geochelone radiata due to overexploitation, ORYX, 2003, Vol:37, Pages:338-343, ISSN:0030-6053(doi)
  • Milner-Gulland EJ, Bukreevea OM, Coulson T, et al, Conservation - Reproductive collapse in saiga antelope harems, NATURE, 2003, Vol:422, Pages:135-135, ISSN:0028-0836(doi)
  • Lindstrom J, Coulson T, Kruuk L, et al, Sex-ratio variation in Soay sheep, BEHAV ECOL SOCIOBIOL, 2002, Vol:53, Pages:25-30, ISSN:0340-5443(doi)
  • Mysterud A, Coulson T, Stenseth NC, The role of males in the dynamics of ungulate populations, J ANIM ECOL, 2002, Vol:71, Pages:907-915, ISSN:0021-8790
  • Clutton-Brock TH, Coulson T, Comparative ungulate dynamics: the devil is in the detail, PHILOS T ROY SOC B, 2002, Vol:357, Pages:1285-1298, ISSN:0962-8436(doi)
  • Carbone C, Christie S, Conforti K, et al, The use of photographic rates to estimate densities of cryptic mammals: response to Jennelle et al., ANIM CONSERV, 2002, Vol:5, Pages:121-123, ISSN:1367-9430(doi)
  • Balmford A, Clegg L, Coulson T, et al, Why conservationists should heed Pokémon., Science, 2002, Vol:295
  • Coulson T, Lindstrom J, Cotgreave P, Ecology - Seeking new recruits, SCIENCE, 2002, Vol:295, Pages:2023-2024, ISSN:0036-8075
  • Clutton-Brock TH, Coulson TN, Milner-Gulland EJ, et al, Sex differences in emigration and mortality affect optimal management of deer populations., Nature, 2002, Vol:415, Pages:633-637, ISSN:0028-0836(doi)
  • Amos W, Wilmer JW, Fullard K, et al, The influence of parental relatedness on reproductive success, P ROY SOC LOND B BIO, 2001, Vol:268, Pages:2021-2027, ISSN:0962-8452
  • Coulson T, Catchpole EA, Albon SD, et al, Age, sex, density, winter weather, and population crashes in Soay sheep, SCIENCE, 2001, Vol:292, Pages:1528-1531, ISSN:0036-8075
  • Milner-Gulland EJ, Shea K, Possingham H, et al, Competing harvesting strategies in a simulated population under uncertainty, ANIM CONSERV, 2001, Vol:4, Pages:157-167, ISSN:1367-9430
  • Coulson T, Mace GM, Hudson E, et al, The use and abuse of population viability analysis, TRENDS ECOL EVOL, 2001, Vol:16, Pages:219-221, ISSN:0169-5347
  • Carbone C, Christie S, Conforti K, et al, The use of photographic rates to estimate densities of tigers and other cryptic mammals, ANIM CONSERV, 2001, Vol:4, Pages:75-79, ISSN:1367-9430
  • Albon SD, Coulson TN, Brown D, et al, Temporal changes in key factors and key age groups influencing the population dynamics of female red deer, J ANIM ECOL, 2000, Vol:69, Pages:1099-1110, ISSN:0021-8790
  • Coulson T, Milner-Gulland EJ, Clutton-Brock T, The relative roles of density and climatic variation on population dynamics and fecundity rates in three contrasting ungulate species, P ROY SOC LOND B BIO, 2000, Vol:267, Pages:1771-1779, ISSN:0962-8452
  • Grenfell BT, Finkenstadt BF, Wilson K, et al, Ecology - Nonlinearity and the Moran effect, NATURE, 2000, Vol:406, Pages:847-847, ISSN:0028-0836
  • Catchpole EA, Morgan BJT, Coulson TN, et al, Factors influencing Soay sheep survival, J R STAT SOC C-APPL, 2000, Vol:49, Pages:453-472, ISSN:0035-9254
  • Milner-Gulland EJ, Coulson TN, Clutton-Brock TH, On harvesting a structured ungulate population, OIKOS, 2000, Vol:88, Pages:592-602, ISSN:0030-1299
  • Paul RE, Coulson TN, Raibaud A, et al, Sex determination in malaria parasites., Science, 2000, Vol:287, Pages:128-131, ISSN:0036-8075
  • Coulson T, Albon S, Slate J, et al, Microsatellite loci reveal sex-dependent responses to inbreeding and outbreeding in red deer calves, EVOLUTION, 1999, Vol:53, Pages:1951-1960, ISSN:0014-3820
  • Stewart GR, Boussinesq M, Coulson T, et al, Onchocerciasis modulates the immune response to mycobacterial antigens, CLIN EXP IMMUNOL, 1999, Vol:117, Pages:517-523, ISSN:0009-9104
  • Coulson T, Albon S, Pilkington J, et al, Small-scale spatial dynamics in a fluctuating ungulate population, J ANIM ECOL, 1999, Vol:68, Pages:658-671, ISSN:0021-8790
  • Grenfell BT, Wilson K, Finkenstadt BF, et al, Noise and determinism in synchronized sheep dynamics, NATURE, 1998, Vol:394, Pages:674-677, ISSN:0028-0836
  • Coulson TN, Albon SD, Pemberton JM, et al, Genotype by environment interactions in winter survival in red deer, J ANIM ECOL, 1998, Vol:67, Pages:434-445, ISSN:0021-8790
  • Coulson TN, Pemberton JM, Albon SD, et al, Microsatellites reveal heterosis in red deer., Proc Biol Sci, 1998, Vol:265, Pages:489-495, ISSN:0962-8452(doi)
  • Coulson T, Albon S, Guinness F, et al, Population substructure, local density, and calf winter survival in red deer (Cervus elaphus), ECOLOGY, 1997, Vol:78, Pages:852-863, ISSN:0012-9658
  • Pemberton JM, Smith JA, Coulson TN, et al, The maintenance of genetic polymorphism in small island populations: large mammals in the Hebrides., Philos Trans R Soc Lond B Biol Sci, 1996, Vol:351, Pages:745-752, ISSN:0962-8436(doi)

    Chapters

    • T.H. Clutton-Brock , B.T. Grenfell , T. Coulson , et al, Population dynamics of Soay sheep, ln:Soay Sheep: Dynamics and Selection in an Island Population, Editor(s): Clutton-Brock, Pemberton, Cambridge, Cambridge University Press, 2004, Pages: 52-112
    • T.H. Clutton-Brock , J.M. Pemberton , T. Coulson , et al, The sheep of St.. Kilda, ln:Soay Sheep: Dynamics and Selection in an Island Population, Editor(s): Clutton-Brock, Pemberton, Cambridge, Cambridge University Press, 2004, Pages: 17-51
    • T.H. Clutton-Brock , T.N. Coulson , Comparative ungulate dynamics: the devil is in the detail, ln:Wildlife Population Growth Rates, Editor(s): Sibly, Hone, Cambridge, Cambridge University Press, 2003, Pages: 249-269
    • T. Coulson , E.J. Hudson , When is the birth rate the key factor associated with population dynamics?, ln:Reproductive Science and Integrated Conservation, Editor(s): Holt, Pickard, Rodger, Wildt, Cambridge, Cambridge University Press, 2003, Pages: 114-130
    • J.M. Pemberton , D.W. Coltman , T.N. Coulson , Using microsatellites to measure the fitness consequences of inbreeding and outbreeding., ln:Microsatellites: Evolution and Applications, Editor(s): Goldstein, Schlotterer, Oxford, Oxford University Press, 1999, Pages: 151-164
    • J.M. Pemberton , J.A. Smith , T.N. Coulson , et al, The maintenance of genetic polymorphism in small island populations: large mammals in the Hebrides, ln:Evolution on Islands, Editor(s): Grant, Cambridge, Cambridge University Press, 1998, Pages: 51-64
    • S.D. Albon , T.N. Coulson , T.H. Clutton-Brock , Demographic constraints in red deer: can the past predict the future?, ln:In Recent Developments in Deer Biology, Editor(s): Milne, Proceedings of the 3rd International Congress on the Biology of Deer, 1997, Pages: 85-95

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